"Dastych (1987) searched his collection of Echiniscus and found males in four species."

William R. Miller, Sandra K. Claxton, Harold, F. Heatwole: Tardigrades of the Australian Antarctic Territories: Males in the Genus Echiniscus (Tardigrada: Heterotardigrada). Zoologischer Anzeiger 238 (1999) p. 304.

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